“…But if we allow that variability may arise in the nervous pathways as well as in the physical stimulus, it becomes evident that the prediction of a square root law from the quantum variability of light must rest on one of the following assumptions: (1) The discriminatory response is determined at the receptor level, immediately on the absorption of quanta-but there is ample evidence, reviewed by Treisman (1966), that the behavioral threshold is more centrally determined. (2) The absorption of a quantum or quanta triggers a sequence of neural events that result in "messages" being transmitted to the central locus at which the discriminatory response is determined, the magnitude of the total message indicating the size of the stimulus; the variability or "noise" in this central transmission is negligible, so that the variation of the central message from trial to trial reflects only the Poisson variability of the stimulus.…”