In the late 1960s and early 1970s, evolutionary biology underwent a scientific revolution in which poorly defined models of evolutionary change were replaced with a theoretically rigorous program of research that integrated the major findings of evolutionary biology, animal behavior, and genetics. This integration included a method, known as the adaptationist program, for identifying and describing design in organisms, a method that matches the engineering requirements inherent in adaptive problems to features in the organism that are designed to solve those problems.In this chapter, I apply the methods of the adaptationist program to human anger. The resultant theory, called the recalibrational theory, states that anger is the output of a cognitive mechanism designed by natural selection to negotiate conflicts of interest. The causes of anger, the behavior it produces, the factors that mitigate it, and its effects on physiology, perception, and cognition can be explained by reference to this adaptive function.In brief, the recalibrational theory states that anger is a system designed by natural selection to recalibrate the weight another individual places on the angry person's interests so that they become less likely to impose costs or deny benefits to the angry individual. Anger deploys two main strategies to convince the target to treat the angry individual better: (a) toward those with