A systematic revision of the family Harpetidae (Trilobita)

Abstract

The systematics of the ten valid harpetid trilobite genera are reviewed. Seven are revised, using standard parsimony and three-item analysis. The monophyly of the Harpetidae is confirmed, and all ingroup genera can be defended as monophyletic groups except for the nonmonophyletic ScotolJarpes group. Emended diagnoses are provided for all the genera within the family. The three subfamilies Dolichoharpinae, Eoharpetinae and Harpetinae are supressed within the Harpetidae. The genera AlIstraloharpes and Sinoharpes are placed in synonymy with DlIbhglasina. Thorslundops and Wegeiinia are placed in synonymy with Hibbertia, and the subgenus FritclJaspis placed in synonymy with LiolJarpes. Reticuloharpes and HelioJwrpes are placed in synonymy with Harpes. The Harpetidae, along with the Entomaspididae and Harpididae, is considered to belong in the Harpetida, which is herein raised to ordinal rank within the subclass Libristoma. Glasgow, u.K.; MBT, Museum of Natural History,

Sheltered Gregarious Behavior of Middle Ordovician Harpetid Trilobites

“…This is a contribution of the Agence Nationale de la Recherche, Programme Blanc SIMI 5-6 RALI. Paleogeographic reconstruction adapted from Cocks and Torsvik (2002) and Manda (2008) with data obtained from a variety of sources (Barrande 1856(Barrande , 1872Whittington 1949;Henry and Phillipot 1968;Romano 1975;Romano and Henry 1982;Kennedy 1988;Koch and Lemke 1995;Ebach and McNamara 2002;Corbacho 2011). ATA 5 Armorican Terrane Assemblage.…”

“…According to the revision by Whittington (1949) and the more recent study of Ebach and McNamara (2002), Eoharpes includes six species including the type species Eoharpes primus and E. benignensis, both from the Barrandian area. Recently, Eoharpes sp.…”

Late Ordovician trilobites from the Xiazhen Formation in Zhuzhai, Jiangxi Province, China

“…Remarks.-Of Hibbertia species (Ebach and McNamara 2002), Hibbertia aodiensis sp. nov. is similar to H. inghami (= Paraharpes inghami) (Owen 1981: pl.…”

Aphyly: A Systematic Designation for a Taxonomic Problem

“…The harpetid trilobite genus Lioharpes was found to be nonmonophyletic as certain members of the genus are more closely related to members of other genera then they are to each other (Ebach and McNamara 2002) (Fig. 3).…”