“…Instead, managers have relied on indirect measures of relative abundance, such as harvest statistics and catch-per-unit-effort, or have conducted extensive radiotelemetry studies to estimate densities (Rausch 1967, Roughton and Sweeny 1982, Fuller and Snow 1988, Burch et al 2005). More recently, transect intercept probability sampling and the sampling unit probability estimator (SUPE), which use the probability of observing track networks in the snow (Becker 1991, Van Sickle and Lindzey 1991, Becker et al 1998, Patterson et al 2004, have been used because they are efficient for extremely large study areas (!10,000 km 2 ), do not rely on collared animals, and provide confidence intervals (Becker et al 1998). However, these approaches are not feasible in areas where track networks cannot be followed continuously from aircraft due to heavy forest cover.…”