“…Recently, there has been increasing empirical and theoretical work on the non-random foraging movements of predators which often responses to prey-contact stimuli such as spatial variation in prey density [10,39], or different type of signals arising directly from prey [74]. See more related examples of mathematical models in [48,44,11,7,12,21,9,43,15,31,27,55]. Kareiva [40] provided a good review on varied mathematical models that deal with dispersal and spatially distributed populations and pointed out the needs of including non-random foraging movements in meta-population models.…”