“…On the basis of numerous examples, Zatoń et al (2016b) rejected this hypothesis and supported the idea that microconchids originated in the Late Ordovician in shallow shelf environments and later colonized marginal marine environments, and eventually freshwater habitats by the Early Devonian through the evolution of osmoregulation (Zatoń et al, 2012a), as did ostracodes during their invasion of freshwater habitats (Bennett, 2008). Indeed, the colonization of freshwater habitats by microconchids is supported by firm paleontological evidence, e.g., their coexistence with such terrestrial or freshwater components as vascular plant remains (Sandberg, 1963; Brower, 1975; Mastalerz, 1996; Zatoń and Mazurek, 2011; Caruso and Tomescu, 2012; Florjan et al, 2012; Zatoń and Peck, 2013; Shcherbakov et al, 2021), charophytes (Ilyes, 1995; Zatoń and Peck, 2013; Shcherbakov et al, 2021), spinicaudatans, and freshwater ostracodes and bivalves (Trueman, 1942; Zatoń and Peck, 2013; Shcherbakov et al, 2021).…”