“…Generally, the contribution of NSC stored before heading for fi nal grain yield is about 30% (Cock and Yoshida, 1972;Akita, 1989). However, much evidence has been accumulated to indicate that NSC accumulated before the onset of grain-fi lling plays a more important role in rice grain-fi lling than merely compensation for a shortage of assimilates during the grain-fi lling (Takeda et al, 1980;Seo and Ota, 1983;Akita, 1989;Nakamura et al, 1992;Sumi et al, 1996;Tsukaguchi et al, 1996; Horie et al, 1997).…”