“…For example, in addition to frequently studied mechanisms for the evolution of cooperation (Nowak, 2006), more realistic aspects, such as detailed psychological mechanisms (Stevens et al, 2011) and higher level of inter-individual variation, e.g. through different types of agent migrations (Fu and Nowak, 2013;Hadzibeganovic et al, 2009;Han et al, 2014;Helbing and Yu, 2009), diversity of reproduction rates and strategyselection time scales (Rong et al, 2013;Wu et al, 2009), varied learning motivation (Zhang et al, 2010), or other random factors (McNamara et al, 2004) need to be included in next model generalizations. Moreover, the influence of individual development (Gottlieb, 2002) and social learning (Laland et al, 2000;Zhang et al, 2012) on behavioral variation and the resulting novel adaptations should more explicitly be addressed in future computational models.…”