“…In total, these four study areas cover 226,553 km 2 of the western Arctic-Boreal region of North America (Figure 1, inset), and span Arctic tundra, boreal forest, peat plateau, lowland wetland, and upland shield terrains. The 33,383 km 2 YFB study area is characterized by fluvially disconnected wetlands surrounding a wide low-gradient reach of the Yukon River in north central Alaska (e.g., Anderson et al, 2013;Pitcher et al, 2019). The 82,200 km 2 MRV study area includes the Mackenzie River Delta, a vast, fluvially connected wetland, the Tuktoyaktuk Peninsula, a polar desert with continuous permafrost and thousands of thermokarst lakes, and boreal forest and plateaus further south along the Mackenzie and Peel Rivers (Burn & Kokelj, 2009;Plug et al, 2008).…”