Climate change is considered a major threat to ocean ecosystems and human well-being (Blanchard et al., 2017; IPBES, 2019;Lotze et al., 2019). Changes in temperature, oxygen and other biogeochemical properties alter species distribution, growth, reproduction, abundance and trophic ecosystem structure (Cheung, Sarmiento, et al., 2013; IPCC, 2019). For example, many commercially important fishes and invertebrates are shifting their distribution towards polar, offshore or deeper waters to remain in their preferred thermal range . In addition, intensifying of marine heatwaves are affecting ecosystem structure and functioning (IPCC, 2019). Furthermore, due to the ongoing uptake of atmospheric CO 2 , the ocean is progressively acidifying (IPCC, 2019) which can be amplified by coastal upwelling (Feely et al., 2008), affecting cellular metabolism, organism physiology and ecosystem-level dynamics (Gattuso & Hansson, 2011).