We examined tracking accuracy and bias (mean-level and projection) in people’s perceptions of their romantic partner’s relationship approach and avoidance motives, similarity in partners’ motives, and positive and negative emotions as potential cues used to make judgments about a partner’s daily motives and motives during shared activities. Using data from 2 studies, 1 using daily diaries (N = 2,158 daily reports), the other using reports of shared activities (N = 1,228 activity reports), we found evidence of tracking accuracy and projection across samples; we also found evidence of mean-level bias such that people underperceived their partner’s approach (daily) and avoidance motives (daily and in shared activities). Partners had similar daily approach and avoidance motives but were not similar in their motives during shared activities. Further, our studies indicated that emotions often serve as relevant, available, and detectable cues for judging a partner’s motives. The results demonstrate that accuracy and bias are both present in judgments of a romantic partner’s approach and avoidance motives, and that people often, but not always, use their partner’s emotions to make such judgments.