InTRoduCTIonThe genus Themeda Forssk. (Gramineae, Andropogoneae) has about 27 species in the (sub)tropical grasslands of the Old World (Clayton et al. 2014). Some giant species (up to 6 m tall) may be vegetation forming and dominate grass wildernesses nearly impenetrable because of their razor sharp leaves, while the pungent calli of the fertile spikelets penetrate clothing and skin. Still, with their huge, pendulous inflorescences they are quite decorative.Hackel (1889: 656) already complained "in nullo certe genere nomina specierum magis confusa quam in Anthistiria [= The meda]" ("Without any doubt [there is] no genus where the names of the species are more confused than in Anthistiria") and this is shown by the impressive synonymy, to which he himself significantly contributed, and was also experienced here.
HISToRyForsskåhl (1775) described the genus with only Themeda tri andra. He derived the generic name from the Yemeni word Thaemed. According to an anonymous source on the internet this would allude to "a depression in which water lies after rain, subsequently drying in summer". The reason for the epithet is not clear, it is quite usual for a grass floret to have three stamens, while, alternatively, there are not three but four reduced spikelets forming an involucre in the partial inflorescence here.This was not the first species that was known to Western science. At least one was already described and depicted by Rumphius (1750): Gramen arguens Rumph., validated by Linnaeus (1762: 117) as Stipa arguens L.Another may be Calamagrostis Rumph. It is not certain whether it is a Themeda at all. Rumphius described it as a giant grass forming tussocks behind which one can hide, or dense shrubbery which can conceal wild animals. He compared it to his Arundo farcta [= Miscanthus floridulus (Labill.) Warb.] and 'siree', i.e. 'sereh', Lemongrass, Cymbopogon citratus (DC.) Stapf, but said it served no purpose at all. Local uses included that as of T. arguens and in decorative wedding arches. The illustration is rather crude, and it is no wonder that Linnaeus (1762: 65) cited it under Schoenus lithospermus L. (= Scleria lithosperma (L.) Sw., Cyperaceae). It was regarded as T. gi gantea by Merrill (1917). This is erroneous, as that species is a Philippine near-endemic. The only giant species of Themeda known to me from Ambon is T. villosa (Poir.) A.Camus var. polyantha (Brongn. ex Buse) Veldk.Another species cultivated in the Uppsala Botanical Garden with unknown provenance was described by Linnaeus (1771) as Andropogon nutans, a combination he already had used in 1753 for an American species, now Sorghastrum nutans (L.) Nash (Veldkamp 1984). In 1774 he renamed this to Andropogon quadrivalvis L.Houttuyn (1782) under Stipa arguens noted a curious grass that he had received from Thunberg collected in Japan, which in no way, he noted, could be the same as Linnaeus's species. He provided a beautiful illustration, but no name. His specimen is in G. Thunberg (1784a, b) described his duplicate in UPS as Andropogon ciliatum Thu...