“…It is increasingly clear that the response of phytoplankton to multiple stressors acting concurrently cannot be obtained by superimposing their separate responses (Boyd and Brown, 2015;Zhu et al, 2016;Boyd et al, 2016;Luxem et al, 2017;Andrew et al, 2019;Strzepek et al, 2019;Trimborn et al, 2019;Boyd, 2019). Satellite observations in the Southern Ocean show complex patterns of environmental change; surface warming in much of the Northern zone contrasts with slight cooling trends over the last 40 years further south (Maheshwari et al, 2013;Kostov et al, 2016;Sallée, 2018); average sea ice concentration has decreased in the Amundsen Sea but increased in parts of the Weddell, Bellingshausen and Ross Seas (Vaughan et al, 2013;Zhang et al, 2018;Pinkerton, 2019;IPCC, 2019); irradiance at the sea-surface has increased north of the Subantarctic Front and generally reduced to the south over the last 20 years; over the same period, mixed-layer depths have likely shallowed in the Northern zone, and both deepened and shallowed in different parts of the Subantarctic and Antarctic zones (Leung et al, 2015;Pinkerton, 2019).…”