“…Almost all the plants in the Arctic are slow-growing perennials that are adapted to survive stressful abiotic limitations such as low mean annual temperature and precipitation, a short growing season, soil moisture extremes, and low soil nutrient availability (Babb and Whitfield, 1977;Chapin et al, 1978Chapin et al, , 1987Ellis, 1980;Billings, 1987). These conditions are not favorable to the metabolic processes of soil microorganisms (Bliss, 1962;Clein and Schimmel, 1995;Mikan et al, 2002); therefore, while total stocks of N and P in soils may be substantial, a large proportion is bound in organic forms not directly available to plants Billings, 1987;Chapin et al, 1995).…”