Tectonic plates are a key feature of Earth's structure, and their behavior and dynamics are fundamental drivers in a wide range of large-scale processes. The operation of plate tectonics, in general, depends intimately on the manner in which lithospheric plates couple to the convecting interior. Current debate centers on whether the transition from rigid lithosphere to flowing asthenosphere relates to increases in temperature or to changes in composition such as the presence of a small amount of melt or an increase in water content below a specified depth. Thus, the manner in which the rigid lithosphere couples to the flowing asthenosphere is currently unclear. Here we present results from laboratory-based torsion experiments on olivine aggregates with and without melt, yielding an improved database describing the crystallographic alignment of olivine grains. We combine this database with a flow model for oceanic upper mantle to predict the structure of the seismic anisotropy beneath ocean basins. Agreement between our model and seismological observations supports the view that the base of the lithosphere is thermally controlled. This model additionally supports the idea that discontinuities in velocity and anisotropy, often assumed to be the base of the lithosphere, are, instead, intralithospheric features reflecting a compositional boundary established at midocean ridges, not a rheological boundary.seismic anisotropy | lithosphere−asthenosphere boundary | upper mantle | geodynamics | crystallographic texture A lthough plate tectonics is the unifying paradigm in the Earth sciences, important questions remain regarding the physical nature of a tectonic plate. A cornerstone of plate tectonic theory states that plates translate across Earth's surface in a relatively rigid and coherent fashion, with deformation largely concentrated at plate boundaries. Restated, the plates are taken to have a high viscosity with a relatively sharp transition to the less viscous convecting mantle beneath. Referring to plates as the lithosphere and to the underlying rock as the asthenosphere [terminology which predates plate tectonic theory (1)] is now common. Partial decoupling of the asthenosphere from the lithosphere appears essential to plate-tectonic-like behavior (2), but whether a change in material properties at the base of the lithosphere arises from increasing temperature (3) or from transitions in melt (4-7) or water content (8, 9) remains unclear, even in the tectonically simple ocean basins.Recent seismological investigations of oceanic upper mantle indicate a change in composition at depths often associated with the lithosphere−asthenosphere boundary (LAB). Receiver-function studies and underside reflections of SS precursors detect a sharp velocity discontinuity (the Gutenberg discontinuity) at a depth of 40 to 150 km (4, 6, 10-12). The sharpness of this discontinuity may indicate the presence of a small amount of melt (4), and the lack of a strong dependence of the discontinuity depth on plate age (6, 10, 13) is roughl...