“…The success of this work will be measured in the number of forthcoming discoveries stimulated by questions left open here. Euglossines hold a special place among corbiculate bees as they are the only group that is not eusocial and are putatively the earliest-diverging branch (e.g., Schultz et al, 1999Schultz et al, , 2001Engel, 2001b;Noll, 2002;Cardinal & Packer, 2007) within a clade that extends at least to the latest Cretaceous (e.g., Engel, 2000Engel, , 2001aEngel, , 2004Ohl & Engel, 2007), albeit available fossils of orchid bees are much younger (e.g., Engel, 1999Engel, , 2014Hinojosa-Díaz & Engel, 2007b;Michez et al, 2012;Dehon et al, in press). These realities coupled with their unique morphology, behavior, and intimate relationship with orchids makes them of paramount interest, particularly the diverse genus Euglossa which encapsulates 55% of the known species.…”