“…Chaining is typically applied to sites with larger-diameter trees, higher tree cover, and degraded understory vegetation [ 121 , 122 ] and creates greater ground disturbance than mastication and cutting, which can in turn increase the density of invasive annual species [ 89 , 115 , 123 , 124 ]. Thus, chained sites are nearly always seeded and inherent soil disturbance associated with chaining is considered necessary to alter seed bed conditions and increase establishment of seeded species [ 95 , 125 , 126 , 127 , 128 , 129 , 130 , 131 , 132 , 133 , 134 , 135 ]. Mastication is also suitable for sites characterized by later stages of woodland development but is followed by seeding only if pretreatment understory conditions are degraded [ 40 , 60 , 89 ].…”