While Teuschestanthes shares characters with modern Ericales and Sapindales, there are significant non-overlapping differences between Teuschestanthes and modern Sapindales (notably, among others, ovule number). Based on available evidence, however, the position of Teuschestanthes is likely as an early offshoot of the stem clade of core Ericales (Ericales sensu stricto). Its relatively unstable floral plan may presage subsequent bilaterality associated with growing selective pressure by advanced pollinators.