“…Because several taxa range beyond these limits, and to facilitate testing of biogeographic hypotheses, we also included distribution records from adjacent R ıo Magdalena and R ıo Atrato basins of northern Colombia (Maldonado-Ocampo et al, 2006, 2008Mojica et al, 2006) and rivers of southern Chiapas and Mexican portions of the R ıo Grijalva-Usumacinta drainage (Schmitter-Soto, 1999;Miller et al, 2005). Distributional data were compiled through literature searches of books and peerreviewed checklists for Panama (Loftin, 1965;Smith and Bermingham, 2005), Costa Rica (Bussing, 2002;Smith and Bermingham, 2005), Nicaragua (Villa, 1982;Smith and Bermingham, 2005;Matamoros et al, 2012a), Honduras (Matamoros et al, 2009, 2012a), El Salvador (McMahan et al, 2013, Guatemala (Kihn-Pineda et al, 2006) and Belize (Greenfield and Thomerson, 1997).…”