“…Despite high intraspecific variability due to environmental factors and sometimes low interspecific variability across species for some traits (Ribeiro da Luz and Crowley, 2010;Rud et al, 2006), Asner (1998) suggests that plant species should be distinguishable remotely due to differences in biochemical and structural properties. Species differences have been detected remotely (Burkholder, 2010;Narumalani et al, 2009;Pinard and Bannari, 2003;Rud et al, 2006;Schmidt and Skidmore, 2001;Yingying et al, 2011) by examining such 58 vegetation characteristics as total chlorophyll (chl), chl a, and chl b, carotenoids, soluble carbon (C), lignin, foliar N and phosphorus (P), leaf water content, compactness of the mesophyll layer, specific leaf area (SLA), and leaf mass per area (LMA) (Asner and Martin, 2011, 2009;Carlson et al, 2007;Castro-Esau et al, 2004).…”