“…On the Shirakawa sandflat, the sediments of the high to mid-tide zone are unstable (Yamada et al, 2007), which would be unsuitable for juveniles of R. philippinarum to survive in (Breber, 2002;Ponurovskii, 2000Ponurovskii, , 2008Yamada et al, 2007). By contrast, the sediment of the low-tide zone is stable and contains more shell fragments (Yamada et al, 2010), which can be used by R. philippinarum juveniles as a base for their byssus thread attachment to prevent dispersal (de Montaudouin, 1997).…”