“…Conversely, the study of the genetic structure at wider scales (the macro scale approach), including both geographical and taxonomic (i.e., speciation) points of view, will usually imply to detect larger spatio-temporal processes and to work with deeper evolutionary timescales (Wang, 2010;Marske, Rahbek, & Nogués-Bravo, 2013). In this sense, the spatial genetic structure within a species at this macro scale will be the result of different historical and contemporary influences such as connectivity across the range of the species and landscape barriers, environmental adaptation, demographic history, climatic events, or even geomorphological events, among others (Hewitt, 2000;Lee & Mitchell-Olds, 2011;Mairal et al, 2017;Nistelberger, Tapper, Coates, McArthur, & Byrne, 2021). Further, if we include the taxonomic perspective in the analysis of genetic structure in a group of closely related species, we will be able to analyse the processes leading to speciation, which also may involve those previously mentioned (Hart, 2011;Fujita, Leaché, Burbrink, McGuire, & Moritz, 2012; see e.g., Zhou et al, 2012;Ren, Mateo, Guisan, Conti, & Salamin, 2018;Zhao, Gugger, Xia, & Li, 2016).…”