15 Across Eurasia and North America, beaver (Castor spp), their dams and their human-built 16 analogues are becoming increasingly common restoration tools to facilitate recovery of streams 17 and wetlands, providing a natural and cost-effective means of restoring dynamic fluvial 18 ecosystems. Although the use of beaver ponds by numerous fish and wildlife species is well 19 documented, debate continues as to the benefits of beaver dams, primarily because dams are 20 perceived as barriers to the movement of fishes, particularly migratory species such as 21 salmonids. In this study, through a series of field experiments, we tested the ability of juvenile 22 salmonids to cross constructed beaver dams (aka beaver dam analogues). Two species, coho 23 salmon (Oncorhynchus kisutch) and steelhead trout (O. mykiss), were tracked using passive 24 integrated transponder tags (PIT tags) as they crossed constructed beaver dams. We found that 25 when we tagged and moved late-summer parr from immediately upstream of the dams to 26 immediately downstream of them, most of them were detected upstream within 36 hours of 27 displacement. By the end of a 21-day field experiment, 91% of the displaced juvenile coho and 28 54% of the juvenile steelhead trout were detected on antennas upstream of the dams while <1% 29 of the coho and 15% of the steelhead trout were detected on antennas in the release pool below 30 the dams. A similar but shorter 4-day pilot experiment with only steelhead trout produced similar 31 results. In contrast, in a non-displacement experiment, juveniles of both species that were 32 captured, tagged and released in a pool 50 m below the dams showed little inclination to move 33 upstream. Finally by measuring hydraulic conditions at the major flowpaths over and around the 3 34 dams, we provide insight into conditions under which juvenile salmonids are able to cross these 35 constructed beaver dams, which should help guide future restoration efforts. 36 38 across riverine landscapes and have altered many natural processes by reducing ecosystem 39 connectivity. In the past five millennia, millions of dams have been constructed by humans, with 40 over two million built in the USA alone [1, 2]. Currently, efforts are underway to remove many 41 of these dams, with the primary objective of restoring stream connectivity, and more specifically, 42 to improve fish passage [3, 4] .
43While the number of dams built by humans is impressive, there are actually fewer dams in 44 North America now than prior to European colonization, albeit of a different size and materials.45 Historic estimates of North American beaver (Castor canadensis) populations range from 60-400 46 million, suggesting that across their 1.5 x10 7 km 2 range, there was anywhere from 10-60 million 47 beaver dams, mostly made of sticks and mud [5][6][7]. In addition, large wood formed millions of 48 jams, dams and other obstructions that dammed and diverted sediment and water across streams, 49 rivers and even entire valleys [8][9][10]. Historic accounts suppo...