“…The structure of flow over dunes whose morphology is in equilibrium with flow has been researched extensively in laboratory experiments (Bennett & Best, 1995;Best, 2005a;Fedele & Garcia, 2001;Kadota & Nezu, 1999;Kwoll et al, 2016;Lefebvre et al, 2016;Maddux et al, 2003;Mazumder et al, 2009;McLean et al, 1999McLean et al, , 1994Nelson et al, 1993;Nelson & Smith, 1989;Venditti, 2007) and simulated using Computational Fluid Dynamics (Chang & Constantinescu, 2013;Grigoriadis et al, 2009;Omidyeganeh & Piomelli, 2011, 2013a, 2013bOmidyeganeh et al, 2013;Stoesser et al, 2008;Xie et al, 2014), which has led to general agreement concerning the nature of the flow field over idealized dune forms, as outlined in conceptual models of flow over dunes (Best, 2005b; Figure 1). However, natural rivers rarely attain a flow morphology equilibrium due to temporal and spatial variations in hydraulic geometry leading to flow decelerations and accelerations, as well as varying discharge on a range of timescales (Aberle et al, 2010;Amsler & Garcia, 1997;Chen et al, 2012;Prent & Hickin, 2001). Consequently, the equilibrium description of flow structure over dunes encapsulated in Figure 1 might apply to natural channels only in rare instances.…”