“…However, some groups of beetles still retain a considerable ecological role and show signs of biological progress (Danks, 1981;Chernov, 1984;Chernov and Makarova, 2008). This communication is based on collections, field observations, and quantitative surveys in various Arctic and Subarctic localities, the results of which were partly reported earlier by the authors (Chernov, 1966(Chernov, , 1973(Chernov, , 1978a(Chernov, , 1978b(Chernov, , 2004Medvedev and Chernov, 1969;Khruleva, 1986, 2011;Khruleva, 1987Khruleva, , 1989Khruleva, , 1991Khruleva, , 1994Khruleva, , 1996Khruleva, , 1999Khruleva, , 2004Khruleva, , 2007Khruleva, , 2009Penev, 1989;Chernov et al, , 2000Chernov et al, , 2001; Korotyaev, 1999, 2012;Makarova et al, 2007;Chernov and Makarova, 2008), and also on other published data not all of which are explicitly mentioned in the text for brevity reasons; mostly the reviews and summary works are cited.…”