Biosystematics of the genus Nathanella Demoulin (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from South India

New Oriental tribe Iscini, new non-dilatognathan species of Notophlebia Peters & Edmunds 1970 and independent origin of Dilatognathus-type mouth apparatus in Atalophlebiinae (Ephemeroptera: Leptophlebiidae)

“…My investigation of Notophlebia (two species described below) and Nathanella (male and female imagos reared from larvae of N. saraswathiae Sivaramakrishnan, Venkataraman & Balasubramanian 1996) has not revealed shared apomorphies of Notophlebia and Nathanella other than the loss of hind wings. Thus, the assumption about their relationship, which was expressed by Peters & Edmunds (1970), has not been supported yet (nor has it been disproven, because the relationships of Nathanella remain unknown).…”

“…Sivaramakrishnan & Peters (1984) described a second species of Notophlebia-N. jobi Sivaramakrishnan & Peters 1984 as imagos and larvae (associated by rearing), and gave the first diagnosis of Notophlebia based on the larval stage. Later, Sivaramakrishnan et al (1996) described true larva of Nathanella, which appeared to be quite different from the larva of Notophlebia.…”