The automated design of Ant Colony Optimization (ACO) algorithms has become increasingly significant, particularly in addressing complex combinatorial optimization problems. Although existing methods have achieved some success, they still face limitations, particularly the high dependency on expert knowledge, pre-solved data, and challenges in interpretability. Genetic Programming (GP), as a proven technology, has shown potential in optimizing the automated design state transition rules of ACO. However, existing research on GP-ACO is insufficient, particularly in terms of experimental validation and systematic evaluation. To address these issues, this study conducts comprehensive experiments to explore several key questions: the generality of GP-ACO on homogeneously distributed maps, the impact of different ACO variants on the learning capabilities of GP-ACO, the effect of 2-opt local search on the learning capabilities of GP-ACO, the enhancement of learning capabilities through the addition of more global information, and the interpretability of GP-ACO. The findings indicate that GP-ACO exhibits robust generality; variations among ACO variants have minimal impact on learning performance; 2-opt local search can somewhat diminish the performance of GP-ACO in the Max-Min Ant System (MMAS); additional global information can significantly enhance the learning capabilities of GP-ACO; and GP-ACO has good interpretability.