“…Paleoenvironmental studies at Chadian hominin sites (faunal composition and structure, dental mesowear and isotopes) concluded to the existence of a diversified vegetation cover, with a mosaic of forests, woodlands, grasslands, up to desert conditions, in close relationship with aquatic/lacustrine areas in northern Chad during the Messinian Schuster et al, 2006;Jacques, 2007;LeFur et al, 2009;Schuster et al, 2009;Blondel et al, 2010;Otero et al, 2010), and open C 4 -vegetations later on during the Pliocene (Brunet et al, 1997;Zazzo et al, 2000;Geraads et al, 2001;Lee-Thorp et al, 2012), in association with large lake occurrences (Schuster et al, 2009). Paleovegetation modeling for the Messinian does not support the existence of mosaic vegetation in northern Chad, but rather a spread of the tropical savanna biome (Pound et al, 2012).…”