Hillslopes are dynamic landscape features that connect ridges to valleys, linking terrestrial and aquatic ecosystems through water, sediment, and nutrient fluxes (Fan et al., 2019;Stieglitz et al., 2003). The degree to which a hillslope parcel is connected to a downstream waterbody is a function of the energy inputs into the parcel and the likelihood for disconnectivity during downstream transport (Najafi et al., 2021). The strength of this connection is a product of tectonic and climatic drivers (Montgomery & Dietrich, 1992; Whittaker, 2012). Recent large-scale analyses of rivers, which act as end-points to the hillslope continuum, disagree whether tectonic or climatic processes control longitudinal river structure (Chen et al., 2019;Seybold et al., 2021). However, similar large-scale analyses of hillslope connectivity are lacking despite being crucial for understanding how landscapes adapt to climate change and other anthropogenic stressors.Structural (or "static") connectivity sets the long-term propensity for downgradient transport whereas functional (or "dynamic") connectivity varies over the short-term and includes the interplay of spatial and temporal energy fluxes (Bracken et al., 2015;Mahoney et al., 2020aMahoney et al., , 2020b. For example, short-term meteorological conditions may characterize the functional hillslope response whereas long-term climatic conditions may set structural landscape evolution trends. To this end, structural connectivity is informed by landscape topographic properties, including elevation, slope, aspect, curvature, and roughness . Further, topography serves as a "tape recorder" of the history of a landscape as tectonic and climatic drivers leave behind features through uplift and downcutting that encode their historic influence on connectivity (Whittaker, 2012).