“…Among them, temperature (T) and relative humidity (RH) are highly related to meteorological conditions, greatly affecting the germination and growth of various airborne fungi inside and outside WCHs. In particular, in the Korean Peninsula with four distinct seasons, when high internal T and RH conditions are created in summer, and simultaneously the air quality inside the space is stagnant, the germination and growth of rot fungi and imperfect fungi, which gain nutrition from organic cultural heritages, cause biological damage to the materials of the heritage structures (Ham et al, 2021;Hong et al, 2018Hong et al, , 2019Lee et al, 2022;Nandika et al, 2023;Priadi et al, 2021Priadi et al, , 2023Yun et al, 2021). For example, in the 1990s, Seosagan-Jang, one of the storage places for Tripitaka in Haeinsa Temple, had poor ventilation, causing high RH during the rainy season every year and continuous mildew growth on the surface of several printing wood blocks stored inside (Park, 2011).…”