The article engages critically with the literature on the relationship between social movements and political parties. It traces the representation crisis in Chile with a dual focus on the meso-institutional supply side of partisan politics and the microfoundational demand side of protest activity (2006–2019). I argue that the dialectical relationship between political parties’ programmatic dealignment and realignment and social actors’ framing politics determined the magnitude, intensity, and ideational content of protest mobilization. Social actors’ perception of their position in polity structure determined the content of their demands. Savvy actors started with a realignment frame in 2006 to push through socioeconomic reforms from within the parameters of the existing system. They, however, afforded an anti-establishment frame with the ‘social outbreak’ in 2019 to weed out the vestiges of Pinochet’s regime. Social forces pushed political parties to reposition their policy programs, reset agenda priorities, and recast their linkages to society. I draw on interview data, archival works, and Observatory of Conflicts–Cumulative Dataset to substantiate my argument.