“…Regarding the features used for discrimination, many studies have used colour, shape (biological morphology) and texture on an individual basis or in combination (Ahmed, Al-Mamun, Bari, Hossain, & Kwan, 2012;Åstrand & Baerveldt, 2002;Gebhardt & Kühbauch, 2007;Nieuwenhuizen, Tang, Hofstee, Müller, & Van Henten, 2007;Pérez, López, Benlloch, & Christensen, 2000;Persson & Åstrand, 2008;Slaughter, Giles, & Downey, 2008;Swain, Nørremark, Jørgensen, Midtiby, & Green, 2011;Zhang, Kodagoda, Ruiz, Katupitiya, & Dissanayake, 2010). These features are intuitive and easy-to-implement but may have limited discrimination ability under ambient lighting conditions.…”