“…Major challenges remain for developing a holistic understanding of IRES. Low regional synchronization of zero flows reveals unknown processes that control intermittence (Snelder et al, 2013), especially the relative influence of natural drivers like climate (Borg Galea et al, 2019; Skoulikidis et al, 2017; Ward, Wondzell, Schmadel, & Herzog, 2020), geology (Lovill et al, 2018; Ward et al, 2018; Whiting & Godsey, 2016), topography (Jensen et al, 2018; Jensen et al, 2019; Prancevic & Kirchner, 2019), and of human factors such as land use and water withdrawals (de Graaf et al, 2019; Dresel et al, 2018; Skoulikidis et al, 2017). It is also unclear how drying alters the estimation of water balance (Cuthbert et al, 2016) and water travel times (Bansah & Ali, 2019; van Meerveld, Kirchner, Vis, Assendelft, & Seibert, 2019).…”