Theories for the origins of agriculture are still debated, with a range of different explanations offered. Computational models can be used to test these theories and explore new hypotheses; Bowles and Choi [Bowles S, Choi J-K (2013) Proc Natl Acad Sci USA 110(22): [8830][8831][8832][8833][8834][8835] have developed one such model. Their model shows the coevolution of farming and farming-friendly property rights, and by including climate variability, replicates the timings for the emergence of these events seen in the archaeological record. Because the processes modeled occurred a long time ago, it can be difficult to justify exact parameter values; hence, we propose a fitting to idealized outcomes (FIO) method to explore the model's parameter space in more detail. We have replicated the model of Bowles and Choi, and used the FIO method to identify complexities and interactions of the model previously unidentified. Our results indicate that the key parameters for the emergence of farming are group structuring, group size, conservatism, and farming-friendly property rights (lending further support to Bowles and Choi's original proposal). We also find that although advantageous, it is not essential that farming productivity be greater than foraging productivity for farming to emerge. In addition, we highlight how model behaviors can be missed when gauging parameter sensitivity via a fix-all-but-one variation approach.Neolithic revolution | simulation | agent-based modeling | fitting to idealized outcomes | property rights A fter around 190,000 y of anatomically modern humans living as hunter-gatherers, societies began to make a transition to agricultural subsistence systems. These transitions are thought to have occurred independently in several regions of the world between 11,500 and 3,500 y before present (yBP) (1), and farming has spread to most of the world from these centers. This transition had major impacts on human demography, evolution, diet, health, culture, technology, and social inequality (2-5). It is generally agreed that there are at least three separate regions around the world where agriculture originated independently: the Fertile Crescent, Mesoamerica, and China (1, 2, 5). There is also growing evidence for additional independent origins of domestication, which include the eastern United States, the Andes, the Central and South America tropics, New Guinea, West Africa, and southern and northern India (5).A range of different explanations for the development of agriculture have been proposed, including some based on affluence and social competition [e.g., via feasting (6)] and others based on stress [e.g., because of population pressure (7) or climate changes (8)]; some propose revolution (e.g., 9), and some suggest changes were made consciously (e.g., 10), whereas others argue changes were made unconsciously (e.g., 11) (1). Whereas the long-term advantages of farming are clear in that it is a strategy that has generally led to increased carrying capacities and greater reproductive successes when...