“…This means that processes that do not involve direct fitness evaluations, such as conformist transmission, which can be adaptive in spatially heterogeneous environments (Henrich and Boyd, 1998; Nakahashi, Wakano, and Henrich, 2012), and when observational errors are common (Henrich and Boyd, 2002), are not going to effectively spread novel adaptive cultural traits by themselves, and can sometimes even lead to population collapse under specific ecological conditions (Whitehead and Richerson, 2009). However, adaptive cultural evolution occurs when a combination of learning strategies is used, such as: (1) conformist learning combined with individual learning (Boyd and Richerson, 1995; Henrich and Boyd, 1998) or payoff‐biased learning (Henrich and Boyd, 2002; Hong, 2022b); (2) “flexible learning” that enables individual learning to be more accurate or less costly (Boyd and Richerson, 1995); (3) “critical social learning,” where individuals engage in social learning first, and if results are unsatisfactory, then deploy individual learning (Enquist, Eriksson, and Ghirlanda, 2007); and (4) “specialized hybrid learning,” in which individuals deploy asocial learning in some contexts and social learning in other contexts (Boyd and Richerson, 1995; Kharratzadeh et al., 2017).…”