“…Applying group-specific correction factors to observed animal groups can compensate for effects of detection bias when estimating abundance and composition. There are several methods for estimating group-specific detection probabilities in aerial surveys, including sightability models (Steinhorst and Samuel, 1989), distance models (Burnham and Anderson, 1984;Buckland et al, 2004), mark-resight models (White, 1996;Skalski et al, 2005), double-observer models (Graham and Bell, 1989), and methods that combine two or more such techniques often referred to as ''hybrid'' models (Quang and Becker, 1997;Buckland et al, 2010;Burt et al, 2014). These techniques all have individual strengths and weaknesses (Griffin et al, 2013).…”