River networks (i.e., dendritic systems of streams within river basins) are hierarchical features in the landscape revealing patterns that recur within and across basins (Frissell et al., 1986). Attributes of individual reaches (i.e., stream segments) within basins depend on physical drivers, such as geologic features, precipitation patterns, channel morphology, and land cover which interact over spatial scales and sequentially along the basin (Allan, 2004;Poff, 1997;Smiley & Dibble, 2005). Basin morphology and hydrology define the intrinsic lengthwise patterns found in river networks, as well as how these patterns are expressed across basins (