“…However, certain segregation of habitat niches is usually observed. It relied on selection of places differing with regard to the following microhabitat factors: density and type of plant cover, distance to water, soil wetness or water level, soil type or richness, litter layer, and quantity of woody debris (Spencer and Pettus 1966, Brown 1967, Hawes 1977, Wrigley et al 1979, Terry 1981, Neet and Hausser 1990, Sheftel 1994, Shvarts and Demin 1994, Dickman 1995, Lee 1995, Castien and Gosalbez 1999. In some cases separation of habitat optima or mutual habitat exclusion has been observed (Hawes 1977, Neet and Hausser 1990, Sheftel 1994.…”