“…Provided no stand replacing disturbances occur in earlier successional stages, the development of unmanaged forests ends decades to centuries later due to stand replacing disturbances, such as fires, storms, avalanches, mud slides, droughts, and insect outbreaks. Although several classifications of the developmental stages of forests have already been proposed (Krotiuk & Bedernichek, 2021), this opinion distinguishes an early (⁓0–100 years), mid (⁓100–200 years)‐, and late (>⁓200 years) developmental stage with the late developmental stage starting after the net ecosystem carbon balance plateaus (Curtis & Gough, 2018) and ending with a stand replacing disturbance. - The original neutrality hypothesis assumed a closed nutrient cycle and acknowledged that open systems may not experience a decline in productivity at maturity (Odum, 1969). Nutrient cycles in forests are characterized by continuous gains and losses and are thus open systems (Galloway et al., 2004).
…”