“…Second, as a forest recovers following major disturbance, it typically changes from an open stand of young saplings, to a closed-canopied, relatively uniform forest, eventually transitioning into a dynamic system of horizontally and vertically diverse structure as trees die, fall over and are replaced 10.3389/ffgc.2022.1070372 by younger trees (Bormann and Likens, 1979;Oliver and Larson, 1996). Forests subject to frequent, non-catastrophic disturbances (e.g., surface fire) maintain old forest structure through different mechanisms, but both types of forest exhibit relatively large trees, decadence and decay, large snags, and spatial heterogeneity in the old-growth stage (Morgan, 1994;Kaufmann et al, 2007;Johnston et al, 2021). These structural characteristics can be used to classify late-successional forest, and in fact, structural definitions are by far the most common approach to classifying old growth (Hilbert and Wiensczyk, 2007).…”