“…Accordingly, other environmental parameters such as nutrient availability become more important for the diversity of the herb layer. A strong link between herbaceous plant diversity and soil parameters associated with the availability of nutrients has indeed been reported in previous studies (Ramovs and Roberts, 2003;Chust et al, 2006;Marks et al, 2008). Nonetheless, it could also be argued that the observed patterns in herbaceous plant diversity represents a state of transition within these plant communities, with different herbaceous species adapting to changes in the climatic conditions at different rates, so that the influence of temperature and precipitation originally observed in the 1963 dataset becoming hidden as a result of the non-equilibrium state within this component of the forest vegetation (Watson, 1980;Shmida and Wilson, 1985;Keeley et al, 2005).…”