“…These latent variables are constructed without reference to environmental measurements, but they can subsequently be compared with actual environmental data if available. To these three well-known types of gradient analysis we add a fourth, constrained ordination, which has its roots in the psychometric literature on multidimensional scaling (Bloxom, 1978;De Leeuw and Heiser, 1980;. Constrained ordination also constructs axes of variation in overall community composition, but does so in such a way as to explicitly optimize the fit to supplied environmental data (Ter Braak, 1986a, 1987c.…”