The Lower Cretaceous Botucatu Formation records the development of widespread dry–aeolian desert sedimentation throughout the Paraná Basin in south‐west Gondwana. To reconstruct the provenance of the aeolian sediment, petrography, granulometric analysis, U‐Pb detrital zircon ages have been determined from along the southern basin margin in Rio Grande do Sul state (southern Brazil) and Uruguay (Tacuarembó region). The dataset reveals a mean composition Qt89F8L3, comprising very fine to medium‐grained quartozose and feldspatho‐quartzose framework. Heavy mineral analysis reveals an overall dominance of zircon, tourmaline and rutile grains (mean ZTR0.84) with sporadic garnet, epidote and pyrolusite occurrences. The detrital zircon U‐Pb ages are dominated by Cambrian to Neoproterozoic (515 to 650 Ma), Tonian to Stenian (900 to 1250 Ma) and Orosirian to Rhyacian (1.8 to 2.2 Ga) material. The detrital zircon dataset demonstrates a significant lateral variation in sediment provenance: Cambrian to Neoproterozoic detrital zircons dominate in the east, while Tonian to Stenian and Orosirian to Rhyacian ages predominate in the west of the study area. Sandstones are quartz‐rich with dominantly durable zircon, tourmaline and rutile heavy mineral suite, with subtle but statistically significant along‐strike differences in heavy mineral populations and detrital mineralogy which are thought to record local sediment input points into the aeolian system. The similar age spectra of Botucatu desert with proximal Paraná Basin units, the predominance of quartzose, and zircon, tourmaline and rutile components, suggests that recycling is the mechanism responsible for the erg feeding.