Mountains are home to a substantial proportion of biological diversity, especially at tropical latitudes (Spehn et al., 2011). The origin of mountain biotas is rather complex, being driven by both geological and biological processes (Rahbek et al., 2019). Mountain lineages originate both from local lowland lineages through niche shift (evolution) and from preadapted lineages through long-distance dispersal (Merckx et al., 2015). However, these two processes, which vary from region to region, are complicated further by uplift and isolation dependent upon geological evolution and past climate changes. Plant and animal species found today in East African Highlands forests often have close relatives in the forests of Central and West Africa or among the separated peaks of East Africa's mountains (Faden, 1983;Mairal et al., 2017;Measey & Tolley, 2011). Within East Africa, montane communities are a distinct region of biotic significance referred