“…31 Ma: Naumann et al, 2013). An Early Cretaceous age, like the one obtained here, is, however, poorly supported by the fossil record, considering that the earliest evidence for an angiosperm dates back to only 136 Ma (Friis et al, 2006) and that Piper is relatively derived among Piperales (Jaramillo and Callejas, 2004). It is likely that the Early Cretaceous age estimated for the divergence of Piper is an overestimation and an inherent artifact in most calibration analyses dealing with early‐diverging lineages, since these are particularly prone to significant changes in extinction/diversification rates (Bromham and Penny, 2003; Friis et al, 2011), life history traits, and other factors that can affect substitution rates and push back the age of a lineage (Kay et al, 2006).…”