“…Despite our knowledge that human behavior is governed to a non-negligible extent by culturally acquired social norms and institutions, there is still a relative lack of models that describe their evolution. Evolutionary models of cultural change (e.g., Boyd & Richerson, 1985) have tended to consider changes in the frequency of culturally transmitted individual-level traits, but only more recently have such models addressed the evolution of enduring higher-order institutional complexes (see, for example, Gavrilets Anderson, & Turchin, 2010;Ostrom, 2005;Shennan, 2009;van der Leeuw & Kohler, 2007). The conditions favoring the enforcement of social norms within communities are captured to some extent by the literature on the evolution of punishment in the evolutionary theory of cooperation (e.g., Axelrod, 1986;Boyd, Gintis, & Bowles, 2010), but it is clear that these models only begin to scratch the surface of the potential complexity of human cultural rule systems (Hill, Barton, & Hurtado, 2009).…”