The living African painted dog, Lycaon pictus (Temminck, 1820), is a successful predator, occupying a wide range of habitats, including short-grass plains, semidesert, bushy savannas, and upland forest. At present, this species is restricted to limited areas in sub-Saharan Africa and classified as endangered species by the International Union for Conservation of Nature (IUCN) (Githiru et al., 2007). The evolutionary origin of Lycaon pictus remains unresolved in spite of several morphologic and molecular studies (Van Valkenburgh and Koepfli, 1993;Tedford et al., 1995; Zrzavy and Rocanková, 2004;Lindblad-Toh et al., 2005). Phylogenetic relationships with the other wolf-like canids, especially the wolf and the dhole, remain unclear, but recent molecular studies only do not support a monophyletic group formed by these three species (Berdeleben et al., 2005).Lycaon-like dogs are well recorded in the early Pleistocene of Europe, Asia, and Africa, being attributed to different taxa depending on the authors (