Abstract. Global climate cooled from the early Eocene hothouse (∼52–50 Ma) to the latest
Eocene (∼34 Ma). At the same time, the tectonic evolution of the Southern Ocean was
characterized by the opening and deepening of circum-Antarctic gateways, which affected both
surface- and deep-ocean circulation. The Tasmanian Gateway played a key role in regulating ocean
throughflow between Australia and Antarctica. Southern Ocean surface currents through and around
the Tasmanian Gateway have left recognizable tracers in the spatiotemporal distribution of
plankton fossils, including organic-walled dinoflagellate cysts. This spatiotemporal distribution
depends on both the physicochemical properties of the water masses and the path of surface-ocean
currents. The extent to which climate and tectonics have influenced the distribution and
composition of surface currents and thus fossil assemblages has, however, remained unclear. In
particular, the contribution of climate change to oceanographic changes, superimposed on long-term
and gradual changes induced by tectonics, is still poorly understood. To disentangle the effects of tectonism and climate in the southwest Pacific Ocean, we target
a climatic deviation from the long-term Eocene cooling trend: the Middle Eocene Climatic Optimum
(MECO; ∼40 Ma). This 500 kyr phase of global warming was unrelated
to regional tectonism, and thus provides a test case to investigate the ocean's physicochemical
response to climate change alone. We reconstruct changes in surface-water circulation and
temperature in and around the Tasmanian Gateway during the MECO through new palynological and
organic geochemical records from the central Tasmanian Gateway (Ocean Drilling Program Site 1170),
the Otway Basin (southeastern Australia), and the Hampden Beach section (New Zealand). Our results
confirm that dinocyst communities track specific surface-ocean currents, yet the variability
within the communities can be driven by superimposed temperature change. Together with published
results from the east of the Tasmanian Gateway, our new results suggest a shift in surface-ocean
circulation during the peak of MECO warmth. Simultaneous with high sea-surface temperatures in
the Tasmanian Gateway area, pollen assemblages indicate warm temperate rainforests with
paratropical elements along the southeastern margin of Australia. Finally, based on new age
constraints, we suggest that a regional southeast Australian transgression might have been
coincident with the MECO.