“…During the Pleistocene (2.58 myBP-11.7 kyBP), the planet experienced repeated cycles of cooler periods with glacial expansion, and warmer interglacial periods. North America was dominated by three expansive ice sheets ~25 kyBP (Dalton et al, 2023): the Laurentide ice sheet east of the Rocky Mountains to the Atlantic Coast of Canada, the Innuitian ice sheet in the far north and the Cordilleran ice sheet west of the Rocky Mountains to the Pacific Coast (Blaise et al, 1990;Dalton et al, 2022;Dyke, 2004;Dyke et al, 2002). Many species were forced to retreat into isolated pockets of suitable habitat known as refugia (Haffer, 1969), with genetic data supporting the presence of multiple northern refugia including Beringia and the Pacific Northwest for plants (Abbott et al, 2000;Abbott & Comes, 2004;Anderson et al, 2006;Brunsfeld et al, 2001;Shafer et al, 2010;Soltis et al, 1997;Thompson & Whitton, 2006;Tremblay & Schoen, 1999), birds (Barrowclough et al, 2004;Lait & Burg, 2013;van Els et al, 2012), and mammals (Brunsfeld et al, 2001;Leonard et al, 2000;Steppan et al, 1999).…”