“…Analysis of the results of this work and those of Voggenreiter (1999), plus studies of numerous natural hybrids on other mountainous islands of the archipelago (GC, H, P, T), and methodical observations (Arango 2015(Arango , 2016a(Arango , 2016b(Arango , 2017(Arango , 2019a(Arango , 2019b(Arango , 2019c(Arango , 2021a(Arango , 2021b(Arango , 2021c(Arango , 2023a have provided essential information to better understand the natural history of F1 hybrids of the genera Aeonium and Greenovia, which I have synthesized into the following eight principles. The principles governing hybridization in the F1 generation in the genera Aeonium and Greenovia have been expressed as principles and not as laws because, in natural sciences, laws are universal, while principles are subject to biological evolution and changes in the natural environment and may have exceptions (Elgin 2003;Martínez & al.…”